Fiddler Four

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Across
  1. 3. Batoid with venomous caudal spine.
  2. 7. Bivalve attached to Spartina roots.
  3. 14. Oily forage fish central to estuarine trophic transfer.
  4. 16. Minimal tidal range phase of lunar cycle.
  5. 18. Migratory curlew with striped crown.
  6. 19. Salt-tolerant woodland behind primary dunes.
  7. 20. Maximum tidal amplitude phase.
  8. 22. Fish-hunting raptor nesting on channel markers.
  9. 25. Opportunistic mesopredator of barrier islands.
  10. 26. Anadromous migrant dependent on horseshoe crab spawning events.
  11. 28. Brackish-water turtle endemic to tidal creeks.
  12. 30. Newly named coastal dolphin species known for strand feeding.
  13. 31. Secretive marsh rallid with a clattering territorial call.
  14. 32. Burrow-dwelling amphibian of sandy uplands.
  15. 34. Coastal piscivore and state seabird of South Carolina.
  16. 36. Semi-aquatic mustelid of tidal systems.
  17. 37. Surface-feeding seabird with elongated lower mandible.
  18. 44. Particulate organic matter forming the base of estuarine food webs.
  19. 45. Pink wader with spatulate bill.
  20. 47. Cryptic flatfish of sandy bottoms.
  21. 48. Marine oscillation driven by gravitational forces.
  22. 49. Bald-headed wading bird federally recovered from endangerment.
Down
  1. 1. Sessile crustacean cemented to hard substrate.
  2. 2. Larid named for its cackling call.
  3. 4. Ancient chelicerate spawning on sandy beaches.
  4. 5. Fine silty substrate exposed at ebb tide.
  5. 6. Aeolian landform shaped by coastal winds.
  6. 8. Echinoid commonly found bleached on beaches.
  7. 9. Marsh-dwelling rallid (abbrev. form).
  8. 10. Spiny echinoderm of tidal pools.
  9. 11. Biogenic shoreline stabilizer composed of living bivalves.
  10. 12. Medium-sized shorebird with bold wing flashes in flight.
  11. 13. Vibrantly colored passerine of maritime scrub.
  12. 15. Narrow channel winding through marsh.
  13. 17. Tidal wetland (generic term).
  14. 20. Intertidal wetland flooded by lunar cycles.
  15. 21. Large-crested seabird with orange bill.
  16. 23. Local colloquialism for anaerobic marsh mud.
  17. 24. Marsh snail grazing on cordgrass epiphytes.
  18. 27. One-clawed signaling crustacean.
  19. 29. Arboreal lizard capable of chromatophore-based color shifts.
  20. 33. Sandy island system buffering mainland marshes.
  21. 35. Small heron of shaded creeks.
  22. 38. Sea turtle nesting on SC barrier islands.
  23. 39. Striped estuarine fish with molariform teeth.
  24. 40. Dominant cordgrass structuring southeastern salt marsh trophic systems.
  25. 41. Venomous pit viper of freshwater wetlands.
  26. 42. Nocturnal pale beach burrower.
  27. 43. Small seabird (generic).
  28. 46. Elusive felid of maritime forest margins.