verna

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Across
  1. 1. Ice river sculpting valleys over millennia
  2. 3. Darwin’s Origin might find a shelf here
  3. 4. Frozen treeless biome with permafrost soils
  4. 6. Lacking subtlety, blunt as human impact
  5. 7. Whole, unfragmented by borders or layers
  6. 10. No longer extant, recorded in scientific logs
  7. 12. Earth’s vent, exemplified by Mount Nyiragongo
  8. 14. Brackish junction where freshwater greets the sea
  9. 17. Prop-rooted tree anchoring tidal zones
  10. 18. Invertebrate negotiating the soil matrix
  11. 21. Wildflower-dotted grassland buzzing with pollinators
  12. 22. Fish synchronizing movement for survival
  13. 24. Boundary beyond which observation ceases. _____ horizon
  14. 26. Grassland with scattered woody sentinels
  15. 27. Rainforest flora forming protective canopies
Down
  1. 2. Fungus and alga united in symbiosis
  2. 5. Constantly in motion, like metabolic pathways
  3. 8. Uppermost layer hosting epiphytes and arboreal fauna
  4. 9. Pauses in ecological succession or volcanic output
  5. 11. Observer of species interactions and ecosystems
  6. 13. Claim lacking empirical support
  7. 15. Wetland stem that surges with water flow
  8. 16. Accumulated insight encoded in books and genes
  9. 19. Interpreter of stratified Earth’s history
  10. 20. Microscopic cargo for angiosperm fertilization
  11. 23. Colonial cnidarian secreting reef architecture
  12. 25. Coastal protrusion reshaping ocean currents